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The UAE called for “restraint and wisdom� and disputed Riyadh’s allegations. But shortly after that, it said it would withdraw its remaining troops in Yemen. It remained unclear whether the separatists it backs will give up the territory they recently took.

The confrontation threatened to open a new front in Yemen's decade-long war, with forces allied against the Iranian-backed Houthi rebels possibly turning their sights on each other in the Arab world's poorest nation.

It also , neighbors on the Arabian Peninsula that increasingly have competed over economic issues and regional politics, particularly in . Tuesday’s airstrikes and ultimatum appeared to be their most serious confrontation in decades.

“I expect a calibrated escalation from both sides. The UAE-backed Southern Transitional Council is likely to respond by consolidating control,� said Mohammed al-Basha, a Yemen expert and founder of the Basha Report, a risk advisory firm.

“At the same time, the flow of weapons from the UAE to the STC is set to be curtailed following the port attack, particularly as Saudi Arabia controls the airspace.�

Airstrike hits Mukalla

A military statement carried by the state-run Saudi Press Agency announced the strikes on Mukalla, which it said came after ships arrived there from Fujairah in the UAE.

“The ships’ crew had disabled tracking devices aboard the vessels, and unloaded a large amount of weapons and combat vehicles in support of the Southern Transitional Council’s forces,� the statement said.

“Considering that the aforementioned weapons constitute an imminent threat, and an escalation that threatens peace and stability, the Coalition Air Force has conducted this morning a limited airstrike that targeted weapons and military vehicles offloaded from the two vessels in Mukalla,� it added.

It wasn't clear if there were any casualties.

The Emirati Foreign Ministry hours later denied it shipped weapons but acknowledged it sent the vehicles “for use by the UAE forces operating in Yemen.� It also claimed Saudi Arabia knew about the shipment ahead of time.

The ministry called for “the highest levels of coordination, restraint and wisdom, taking into account the existing security challenges and threats.�

The Emirati Defense Ministry later said it would withdraw its remaining troops from Yemen over “recent developments and their potential repercussions on the safety and effectiveness of counter-terrorism operations.� It gave no timeline for the withdrawal. .

Yemen’s anti-Houthi forces not aligned with the separatists declared a state of emergency Tuesday and ended their cooperation with the UAE. They issued a 72-hour ban on border crossings in territory they hold, as well as entries to airports and seaports, except those allowed by Saudi Arabia. It remained unclear whether that coalition, governed under the umbrella of Yemen's Presidential Leadership Council, would remain intact.

The Southern Transitional Council’s AIC satellite news channel aired footage of the strike's aftermath but avoided showing damage to the armored vehicles.

“This unjustified escalation against ports and civilian infrastructure will only strengthen popular demands for decisive action and the declaration of a South Arabian state,� the channel said.

The attack likely targeted a ship identified as the Greenland, a vessel flagged out of St. Kitts. Tracking data analyzed by the AP showed the vessel had been in Fujairah on Dec. 22 and arrived in Mukalla on Sunday. The second vessel could not be immediately identified.

Jens Laerke, a spokesperson for the U.N. humanitarian office, urged combatants to protect civilians and civilian infrastructure, like the port, saying any disruption to its operations “risks affecting the already dire humanitarian situation and humanitarian supply chains.�

Strike comes as separatists advance

Mukalla is in Yemen's Hadramout governorate, which the council seized in recent days. The port city is some 480 kilometers (300 miles) northeast of Aden, which has been the seat of power for anti-Houthi forces after the rebels seized the capital, Sanaa, in 2014.

Yemen, on the southern edge of the Arabian Peninsula off East Africa, borders the Red Sea and the Gulf of Aden. The war there has killed more than 150,000 people, including fighters and civilians, and created one of the world's worst humanitarian disasters.

The Houthis, meanwhile, have launched attacks on hundreds of ships in the Red Sea corridor over the Israel-Hamas war in the Gaza Strip, disrupting . The U.S., which earlier praised Saudi-Emirati efforts to end the crisis over the separatists, has launched airstrikes against the rebels under both Presidents and .

Tuesday's strike in Mukalla comes after Saudi Arabia targeted the council in airstrikes Friday that analysts described as a warning for the separatists to halt their advance and leave the governorates of Hadramout and Mahra.

The council had pushed out forces there affiliated with the Saudi-backed ºÃÉ«tv Shield Forces, another group in the anti-Houthi coalition.

Those aligned with the council have increasingly flown the flag of South Yemen, which was a separate country from 1967-1990. Demonstrators have been rallying to support political forces calling for South Yemen to secede again.

A statement Tuesday from Saudi Arabia's Foreign Ministry directly linked the council's advance to the Emiratis for the first time.

“The kingdom notes that the steps taken by the sisterly United Arab Emirates are extremely dangerous,� it said.

Allies of the council later issued a statement in which they showed no sign of backing down.

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Associated Press writer Jamey Keaten in Geneva contributed to this report.